¹School of Pharmacy, Shaanxi University of Chinese Medicine, Xi’an 712046, China
²Genome Analysis Laboratory of the Ministry of Agriculture, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural Sciences, Shenzhen 518000, China
³Marine and Agricultural Biotechnology Laboratory, College of Geography and Oceanography, Minjiang University, Fuzhou 350108, China
⁴Department of Resources Science of Traditional Chinese Medicines, School of Traditional Chinese Pharmacy, China Pharmaceutical University, Nanjing 210009, China
⁵Shaanxi Innovative Drug Research Center, Shaanxi University of Chinese Medicine, Xi’an 712046, China
⁶School of Bioscience and Engineering, Shaanxi University of Technology, Hanzhong 723000, China
⁷Tibetan Medicine Research Center of Qinghai University, Tibetan Medical College of Qinghai University, Xining 810016,China
*Corresponding author. E-mail: yupingtang9@126.com,zhangxingtan@caas.cn,zhaoyucheng1986@126.com
^†Dingqiao Xu,Ziqi Ye and Yongji Huang contributed equally to the study.