¹Laboratory for Research and Utilization of Qinghai Tibet Plateau Germplasm Resources, Key Laboratory of Qinghai-Tibet Plateau Biotechnology Ministry of Education, Academy of Agriculture and Forestry Sciences, Qinghai University, Xining 810016, China
²National Key Laboratory for Tropical Crop Breeding, College of breeding and multiplication, Sanya Institute of Breeding and Multiplication, Hainan University, Sanya 572025, China
*Corresponding author. E-mail: feichen@hainanu.edu.cn,qhwf324@163.com
^†Both authors contributed equally to the study.