¹State Key Laboratory of Herbage Improvement and Grassland Agro-ecosystems, College of Ecology, Lanzhou University, Lanzhou 730000, China
²College of Life Sciences, Shanxi Normal University, Taiyuan 030031, China
³College of Life Sciences, Northwest Normal University, Lanzhou 730070, China
⁴National Key Laboratory for Tropical Crop Breeding, Sanya Institute of Breeding and Multiplication, Hainan University, Sanya 572025, China
⁵School of Tropical Agriculture and Forestry, Hainan University, Haikou 570228, China
⁶State Key Laboratory of Plant Diversity and Specialty Crops, Institute of Botany, the Chinese Academy of Sciences, Beijing 100093, China
⁷State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China
⁸University of Chinese Academy of Sciences, Beijing 100049, China
*Corresponding author. E-mail: wusd@lzu.edu.cn,feichen@hainanu.edu.cn,jiaoyn@ibcas.ac.cn,liujq@nwipb.ac.cn
^†Both authors contributed equally to the study.