¹Hebei Key Laboratory of Horticultural Germplasm Excavation and Innovative Utilization, Hebei Higher Institute Application Technology Research and Development Center of Horticultural Plant Biological Breeding, College of Horticulture Science and Technology, Hebei Normal University of Science and Technology, Qinhuangdao 066004, China
²School of Life Sciences/School of Basic Medical Sciences, North China University of Science and Technology, Tangshan, Hebei 063210, China
³State Key Laboratories of Agrobiotechnology, Joint International Research Laboratory of Crop Molecular Breeding, Beijing Key Laboratory of Growth and Developmental Regulation for Protected Vegetable Crops, Department of Vegetable Sciences, China Agricultural University, Beijing 100193, China
*Corresponding author. E-mail: zhxiaolan@cau.edu.cn,yanliying0665@hevttc.edu.cn,songxm@ncst.edu.cn
^†Both authors contributed equally to the study.