¹Tangshan Key Laboratory of Cruciferous Vegetable Genetics and Breeding, Hebei Key Laboratory of Plant Biotechnology Research and Application, Faculty of Life Science, Tangshan Normal University, Tangshan, Hebei 063000, China
²School of Life Sciences/School of Basic Medical Sciences/Library, North China University of Science and Technology, Tangshan, Hebei 063210, China
³Faculty of Plantation and Agrotechnology, Universiti Teknologi MARA, Jasin Campus,, Merlimau, Melaka 77300, Malaysia
⁴Beijing Zhongnong Futong Horticulture Co., LTD, Beijing 100029, China
*Corresponding author. E-mail: liqiang@tstc.edu.cn;,475544511@qq.com,suyanbin_223@126.com,songxm@ncst.edu.cn
^†Qiang Li,Yanhong Fu and Chenhao Zhang contributed equally to the study.