¹College of Life Science/Hebei Basic Science Center for Biotic Interactions, Institute of Life Science and Green Development, Hebei University, No. 180 Wusi Dong Road, Baoding 071002, China
²Beijing Key Laboratory of Environment Friendly Management on Fruit Diseases and Pests in North China, Institute of Plant Protection, Beijing Academy of Agriculture and Forestry Sciences, No. 9 dawn Garden Road, Beijing 100081, China
³Plant Protection Institute, Hebei Academy of Agriculture and Forestry Sciences, No. 437 Dongguan Road, Baoding 071000, Hebei, China
⁴College of Life Sciences, Shandong Agricultural University, No. 61 Daizong Street, Tai’an 271018, Shandong, China
*Corresponding author. E-mail: sunjh@hbu.edu.cn,zwkang@hbu.edu.cn
^†Zichao Song,Cheng Qu and Ziying Liu contributed equally to the study.