¹National Key Laboratory for Tropical Crop Breeding, College of breeding and multiplication, Sanya Institute of Breeding and Multiplication, Hainan University, Sanya 572025, China
²College of Tropical Agriculture and Forestry, Hainan University, Danzhou 571700, China
³Department of Biology, Saint Louis University, St. Louis, MO 63103, USA
⁴College of Agricultural & Biotechnology, Zhejiang University, Hangzhou 310085, China
⁵Tea Research Institute, Chinese Academy of Agricultural Sciences, Hangzhou 310008, China
⁶College of Horticulture, Nanjing Agricultural University, Nanjing 210095, China
⁷Hainan Fodu Lianyuan Ecological Agriculture Co. Ltd., Haikou 570105, China
⁸Agronomy Department of Superior School Engineering, University of Almería, 04120 Almeria, Spain
⁹Department of Plant Biotechnology and Bioinformatics, Ghent University, Ghent 9052, Belgium
¹⁰Center for Plant Systems Biology, VIB, Ghent 9052, Belgium
¹¹Department of Biochemistry, Genetics and Microbiology, Centre for Microbial Ecology and Genomics, University of Pretoria, Pretoria 0028, South Africa
¹²College of Horticulture, Academy for Advanced Interdisciplinary Studies, Nanjing Agricultural University, Nanjing 210095, Nanjing, China
¹³National Key Laboratory for Tropical Crop Breeding, Laboratory of Crop Gene Resources and Germplasm Enhancement in South China, Ministry of Agriculture and Rural Affairs, Key Laboratory of Tropical Crops Germplasm Resources Genetic Improvement and Innovation of Hainan Province, Tropical Crops Genetic Resources Institute, Chinese Academy of Tropical Agricultural Sciences, Haikou 571101, China
¹⁴Co-first authors
*Corresponding author. E-mail: feichen@hainanu.edu.cn
^†Junyu Zhang contributed equally to the study.