¹Tea Science Research Institute, Ministry of Agriculture and Rural Affairs Key Laboratory of Biology and Germplasm Enhancement of Horticultural Crops in East China, College of Horticulture, Nanjing Agricultural University, 1 Weigang, Nanjing 210095, China
²State Key Laboratory of Crop Genetics & Germplasm Enhancement and Utilization, College of Horticulture, Nanjing Agricultural University, 1 Weigang, Nanjing 210095, China
³Faculty of Agronomy, Jilin Agricultural University, 288 Newtown Street, Changchun, China
⁴Department of Biology, University of British Columbia, 3333 University Way, Okanagan, Kelowna, BC, V1V 1V7, Canada
⁵Biotechnology Research Institute, Shanghai Academy of Agricultural Science, 2901 Beidi Road, Shanghai 201106, China
*Corresponding author. E-mail: xiongaisheng@njau.edu.cn,jian.zhang@ubc.ca,yaoquanhong_sh@aliyun.com,zhuangjing@njau.edu.cn